Eukaryoten und Prokaryoten sind die beiden Superkingdoms der Organismen. Das eukaryotische Chromosom ist eineindeutig, d. h. es enth�lt eine Doppelhelix der DNA. Die um Nukleosomen gewickelte DNA muss f�r die Bildung des Pr�initiationskomplexes mit seiner RNA-Polymerase und allgemeinen Transkriptionsfaktoren zur Verf�gung stehen. Es scheint, dass die um Nukleosomen gewickelte DNA f�r die Bildung des Pr�initiationskomplexes oft nicht zug�nglich ist, aber f�r die Erkennung durch transkriptionsaktivierende Proteine, auch spezifische Transkriptionsfaktoren genannt, verf�gbar ist.

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